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Normally you can't learn abilities till your combat level reaches the required level but with token upgrades you can unlock the abilities earlier. This will replace said ability with the higher tier automatically and if lycanthropy is too low, the ability will be grayed out and you will be unable to use it. Plan accordingly and don't rush into combat after training.
It should also be noted that upon unlocking claw 5 from the Altar of Norala as a lycan that was not yet lvl 58 (req lvl for claw 5), claw 4 was removed from hotbar AND abilities listed under lycanthropy. Claw 1, Claw 2, claw 3, and Claw 5 were listed but not Claw 4 under abilities. Bug report was submitted and awaiting feedback. This effectively reduced dps noticeably as a main lycan as player was forced to use claw 3. - Volshan 4Aug2020